This is the "Cost Each" multiplied by "Total Qty needed for this shipment" This calculated cost will be the cost for this line.
If you change the "Cost Each" or the "Total Qty needed for this shipment" this value will update however the "Total Costing" will be set to 0.
Initially the "total Costing" field is the value of all the "Calc Cost" added together. If you make a change the "Total Costing" field will be set to blank.
The "Total Costing" field is only a visual of the costs added together. The "Total Costing" field is re-calculated when you "Commit" the order to costing.